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- Robber flies of Germany -
[Key]      Catalogue      Comparisons      Notes      Terminology     

key

 -   Subfamilies
 -   Asilinae
    Machimus-group
 -   M. arthriticus
 -   M. chrysitis
 -   M. cyanopus
 -   M. gonatistes
 -   M. rusticus
 -   M. setibarbus
 -   N. setosulus
 -   T. atricapillus
 -   T. atripes
 -   T. cingulatus
 -   T. cowini
 -   T. micans
 -   T. pyragra
 -   T. strandi

 

 

 

  contents & layout
  & copyright:
  Fritz Geller-Grimm
  Imprint

 
Identification key to the species of the Machimus-group

back to the key to Asilinae start of the key

Currently, there are 14 species of the Machimus-group known from Germany, which are placed in Tolmerus, Machimus, and Neoepitriptus. However, the placement of species is difficult and unsatisfactory due to the lack of well-corroborated generic diagnoses. LEHR (1996) tried to clarify the status of the genera of the Machimina-group, but without success. He has also synonymised Epitriptus with Tolmerus. It is probable that some additional species, especially in the poecilogaster-group, will eventually be found to occur in Germany. Single female specimens can often not be determined without knowledge of males.
The following tables for comparison of taxa are available:


1 a. Front femur with distinct stout bristles ventrally [Fig. 1]
. . . . . . . . . . . . to 2
Fig. 1: Neoepitriptus setosulus: Vorderfemur
1 b. Front femur without distinct stout bristles ventrally and only with hairs [Fig. 2]
. . . . . . . . . . . . to 3
Fig. 2: Machimus: Vorderfemur

2 a. Femora entirely black [Fig. 3]
. . . . . . . . . . . . Machimus arthriticus (Zeller, 1840)
Fig. 3: Machimus arthriticus: lateral
2 b. Femora yellow posteriorly [Fig. 4]
Fig. 4: Neoepitriptus setosulus: lateral

3 a. Legs entirely black (sometimes `knees` paler) [Fig. 5], [Fig. 6]
. . . . . . . . . . . . to 4
Fig. 5: Tolmerus atripes: lateral   Fig. 6: Machimus cyanopus: lateral
3 b. Legs at least in part red or yellow [Fig. 7], [Fig. 8]
. . . . . . . . . . . . to 6
Fig. 7: Machimus rusticus: lateral   Fig. 8: Tolmerus cingulatus: lateral

4 a. Species generally smaller than 18 mm [Fig. 9]; 2-4 scutellar marginal bristles [Fig. 10]
. . . . . . . . . . . . Tolmerus atripes Loew, 1854
Fig. 9: Tolmerus atripes: dorsal   Fig. 10: Tolmerus atripes: Scutellum
4 b. Species generally larger than 18 mm [Fig. 11]; 4-8 scutellar marginal bristles [Fig. 12]
. . . . . . . . . . . . zu 5
Fig. 11: Machimus setibarbus: dorsal   Fig. 12: Machimus cyanopus: Scutellum

5 a. Face beard (mystax) with distinctly stout bristles (especially ventrally) [Fig. 13], [Fig. 24]
. . . . . . . . . . . . Machimus setibarbus (Loew, 1849)
or Machimus intermedius Holmgren in Zetterstedt, 1852
Fig. 13: Machimus setibarbus: Kopf   Fig. 24: Machimus setibarbus: Knebelbart
5 b. Face beard (mystax) with only `normal` bristles [Fig. 15], [Fig. 16]
. . . . . . . . . . . . Machimus cyanopus (Loew, 1849)
Fig. 15: Machimus cyanopus: Kopf   Fig. 16: Machimus cyanopus: Knebelbart

6 a. Femora black, tibiae predominantly black and only red or yellow basally [Fig. 17], [Fig. 18]
. . . . . . . . . . . . to 7
Fig. 17: Machimus gonatistes: lateral   Fig. 18: Machimus rusticus: lateral
6 b. Femora in part red or yellowh [Fig. 19], [Fig. 20]
. . . . . . . . . . . . to 8
Fig. 19: Machimus chrysitis: lateral   Fig. 20: Tolmerus cingulatus: lateral

7 a. Only black bristles on legs; scutum predominantly with black bristles; postpedicel nearly equal in length to first two antennal segments combined [Fig. 21]; eepandria apically slanted [Fig. 22]
. . . . . . . . . . . . Machimus gonatistes (Zeller, 1840)
Fig. 21: Machimus gonatistes: Antenna   Fig. 22: Machimus gonatistes: Hypopygium lateral
7 b. Black and yellow bristles on legs; scutum predominantly with yellow bristles; postpedicel longer than first two antennal segments combined [Fig. 23]; epandria narrow, rounded apically [Fig. 24]
. . . . . . . . . . . . Machimus rusticus (Meigen, 1820)
Fig. 23: Machimus rusticus: Antenna   Fig. 24: Machimus rusticus: Hypopygium lateral

8 a. Species generally larger than 18 mm; 4-8 scutellar marginal bristles [Fig. 25], [Fig. 26]
. . . . . . . . . . . . Machimus chrysitis (Meigen, 1820)
Fig. 25: Machimus chrysitis: lateral   Fig. 26: Machimus chrysitis: Scutellum
8 b. Species generally shorter than 18 mm; 2-4 scutellar marginal bristles [Fig. 27], [Fig. 28]
. . . . . . . . . . . . to 9
Fig. 27: Tolmerus cingulatus: lateral   Fig. 28: Tolmerus cingulatus: Scutellum

9 a. Tibiae red, black apically and with a black ring medially [Fig. 29], [Fig. 30]
. . . . . . . . . . . . to 10
Fig. 29: Tolmerus cingulatus: Tibia   Fig. 30: Tolmerus cingulatus: Tibia
9 b. Tibiae red, but without such a black ring medially [Fig. 31], [Fig. 32]
. . . . . . . . . . . . to 11
Fig. 31: Tolmerus strandi: Beine   Fig. 32: Tolmerus pyragra: Head & front leg

10 a. Hind femora, in posterior view, red dorsally [Fig. 33] - female specimens cannot be assigned properly to species; gonostylus (dististylus) smaller and not strongly angled basally [Fig. 34] - cf. table for comparison!
Fig. 33: Tolmerus cingulatus: Hind femora - posterior   Fig. 34: Tolmerus cingulatus: Gonostylus
10 b. Hind femora, in posterior view, predominantly black dorsally, only base and tip red [Fig. 35] - this character is usually found in males, but does not work well for females, which are red dorsally; gonostylus (dististylus) broader and distinctly angled at base [Fig. 36]
. . . . . . . . . . . . Tolmerus cowini (Hobby, 1946)
Fig. 35: Tolmerus cowini: Hind femora - posterior   Fig. 36: Tolmerus cowini: Gonostylus

11 a.
 
males (cf. table for comparison)
. . . . . . . . . . . . to 12

11 b.
 
females (female specimens cannot be assigned to species without males, cf. table for comparison)
. . . . . . . . . . . . to 15

 

 

 

 

 

 

 


12 a. Last sternite with a long, bifid projection posteriorly [Fig. 37], [Fig. 38]
Fig. 37: Tolmerus atricapillus: Hypopygium lateral   Fig. 38: Tolmerus atricapillus: letztes Sternit ventral
12 b. Last sternite with a usual, more or less straight hind margin [Fig. 39]
. . . . . . . . . . . . to 13
Fig. 39: Tolmerus pyragra: Hypopygium lateral

13 a. Gonostylus excavated apically [Fig. 40]; aedeagus short and straight towards tip [Fig. 41]
. . . . . . . . . . . . Tolmerus pyragra (Zeller, 1840)
Fig. 40: Tolmerus pyragra: Gonostylus   Fig. 41: Tolmerus pyragra: Aedeagus
13 b. Gonostylus only weakly excavated apically [Fig. 40]; aedeagus long and bent [Fig. 41]
. . . . . . . . . . . . to 14
Fig. 40: Tolmerus pyragra: Gonostylus   Fig. 41: Tolmerus pyragra: Aedeagus

14 a. Gonostylus short and wide apically [Fig. 42]; aedeagal apodeme small [Fig. 43]
. . . . . . . . . . . . Tolmerus strandi (Duda, 1940)
Fig. 42: Tolmerus pyragra: Gonostylus   Fig. 43: Tolmerus pyragra: Aedeagus
14 b. Gonostylus long and narrower apically [Fig. 44]; aedeagal apodeme larger [Fig. 45]
. . . . . . . . . . . . Tolmerus micans (Meigen, 1820)
Fig. 44: Tolmerus pyragra: Gonostylus   Fig. 45: Tolmerus pyragra: Aedeagus

15 a. Ovipositor cylindrical in cross-section (not distinctly laterally compressed) [Fig. 46], [Fig. 47]
. . . . . . . . . . . . Tolmerus strandi (Duda, 1940)
Fig. 46: Tolmerus strandi: Ovipisitor   Fig. 47: Tolmerus strandi: Ovipisitor
15 b. Ovipositor laterally compressed [Fig. 48], [Fig. 49]
. . . . . . . . . . . . to 16
Fig. 48: Tolmerus atricapillus: Ovipositor   Fig. 49: Tolmerus pyragra: Ovipositor

16 a. [no assignment to species possible without males]

. . . . . . . . . . . . Tolmerus micans (Meigen, 1820)

. . . . . . . . . . . . Tolmerus pyragra (Zeller, 1840)

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

 

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